Publications (recent)

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SDSU undergraduates, SDSU Graduate Students, SDSU Postdocs, Dr. Hedin


Hedin M, Milne MA. 2023. New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains. ZooKeys 1145: 1-130.


Azevedo GHF, T Bougie, M Carboni, M Hedin, MJ Ramírez.. 2022. Convergence, hemiplasy and correlated evolution shape morphological diversity related to web-less lifestyle in the two-clawed spiders. Insect Systematics & Diversity 6(5), 1-14.

Derkarabetian S, J Starrett, M Hedin. 2022. Using natural history to guide supervised machine learning for cryptic species delimitation with genetic data. Front Zool 19: 8.

Ciaccio E, A Debray, M Hedin. 2022. Phylogenomics of paleoendemic lampshade spiders (Araneae, Hypochilidae, Hypochilus), with the description of a new species from montane California. ZooKeys 1086: 163-204.

Derkarabetian S, P Paquin, J Reddell, M Hedin. 2022. Conservation genomics of federally endangered Texella harvester species (Arachnida, Opiliones, Phalangodidae) from cave and karst habitats of central Texas. Conservation Genetics,

Azevedo GHF, T Bougie, M Carboni, M Hedin, MJ Ramírez.. 2022. Combining genomic, phenotypic and Sanger sequencing data to elucidate the phylogeny of the two-clawed spiders (Dionycha). Molecular Phylogenetics and Evolution 166, 107327.


Bougie TC, A Brelsford, M Hedin. 2021. Evolutionary impacts of introgressive hybridization in a rapidly evolving group of jumping spiders (F. Salticidae, Habronattus americanus group), Molecular Phylogenetics and Evolution 161, 107165,

Derkarabetian S, CM Baker, M Hedin, CE Prieto & G Giribet. 2021. Phylogenomic reevaluation of Triaenonychoidea (Opiliones : Laniatores), and systematics of Triaenonychidae, including new families, genera, and species. Invertebrate Systematics 35, 133–157.

Ledford J, S Derkarabetian, C Griswold, C Ribera & M Hedin. 2021. Phylogenomics and Holarctic biogeography of leptonetid spiders (Araneae, Leptonetidae). Invertebrate Systematics 35 (3), 332-349.

Girard MD, DO Elias, G Azevedo, K Bi, MM Kasumovic, JM Waldock, EB Rosenblum & M Hedin. 2021. Molecular phylogenomics of peacock spiders and their kin (Maratus, Euophryinae, Salticidae), with implications for the evolution of male courtship displays. Biol J Linn Society,

Ramírez MJ, ILF Magalhaes, S Derkarabetian, J Ledford, CE Griswold, HM Wood & M Hedin 2021. Sequence-capture phylogenomics of true spiders reveals convergent evolution of respiratory systems. Systematic Biology 70 (1), 14–20.

Rivera C, M Hedin, AC Mason, WP Maddison & DO Elias. 2021 Complex courtship in the Habronattus clypeatus group (Araneae, Salticidae). The Journal of Arachnology 48 (3), 221-232.


Opatova V, CA Hamilton, M Hedin, L Montes de Oca, J Král, CR Haddad & JE Bond. 2020. Systematics and classification of the spider infraorder Mygalomorphae using genomic scale data. Systematic Biology 69, 671–707.

Maddison WP, DR Maddison, S Derkarabetian & M Hedin. 2020. Sitticine jumping spiders: phylogeny, classification and chromosomes (Araneae: Salticidae: Sitticini). ZooKeys 925, 1-54.

Hedin M, S Foldi & B Rajah-Boyer. 2020. Evolutionary divergences mirror Pleistocene paleodrainages in a rapidly-evolving complex of oasis-dwelling jumping spiders (Salticidae, Habronattus tarsalis). Molecular Phylogenetics and Evolution 144, 106696.


Derkarabetian S, S Castillo, PK Koo, S Ovchinnikov & M Hedin. 2019.
A demonstration of unsupervised machine learning in species delimitation,
Molecular Phylogenetics and Evolution, Volume 139, 106562.

Hedin M, S Derkarabetian, A Alfaro, MJ Ramírez & JE Bond. 2019. Phylogenomic analysis and revised classification of atypoid mygalomorph spiders (Araneae, Mygalomorphae), with notes on arachnid ultraconserved element loci. PeerJ, 7, e6864.


Hedin M, S Derkarabetian, J Blair & P Paquin. 2018. Sequence capture phylogenomics of eyeless Cicurina spiders (Araneae, Hahniidae) from Texas caves, with emphasis on US federally endangered species from Bexar County. Zookeys, 769:49-76.

Derkarabetian S, J Starrett, N Tsurusaki, D Ubick, S Castillo & M Hedin. 2018. A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements. Zookeys, 760:1-36.

Hedin MS Derkarabetian, M Ramírez, C Vink & J Bond. 2018. Phylogenomic reclassification of the world’s most venomous spiders (Mygalomorphae, Atracinae), with implications for venom evolution. Scientific Reports (2018) 8:1636 DOI:10.1038/s41598-018-19946-2

Burns M, M Hedin, N Tsurusaki. 2018. Population genomics and geographical parthenogenesis in Japanese harvestmen (Opiliones, Sclerosomatidae, Leiobunum). Ecol. Evol. 8, 36-52.

Starrett J, C Hayashi, S Derkarabetian & M Hedin. 2018. Cryptic elevational zonation in trapdoor spiders (Araneae, Antrodiaetidae, Aliatypus janus complex) from the California southern Sierra Nevada. Molecular Phylogenetics & Evolution, 118, 403-413.


Carlson DE & M Hedin.. 2017. Comparative transcriptomics of Entelegyne spiders (Araneae, Entelegynae), with emphasis on molecular evolution of orphan genes. PLoS One 12 (4), e0174102.

Hedin M & M McCormack. 2017. Biogeographic evidence for common vicariance and rare dispersal in a southern Appalachian harvestmen (Sabaconidae, Sabacon cavicolens). Journal of Biogeography 44 (7), 1665-1678. DOI: 10.1111/jbi.12973.

Burns M, J Starrett, S Derkarabetian, CH Richart, A Cabrero & M Hedin.. 2017. Comparative performance of double-digest RAD sequencing across divergent arachnid lineages. Molecular Ecology Resources 17 (3), 418-430. DOI: 10.1111/1755-0998.12575

Starrett J, S Derkarabetian, M Hedin, RW Bryson Jr., JE McCormack, & BC Faircloth. 2017. High phylogenetic utility of an Ultraconserved element probe set designed for Arachnida. Molecular Ecology Resources 17 (4), 812-823. DOI: 10.1111/1755-0998.12621

Wheeler WC, 17 others,  M Hedin, 16 others. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sample. Cladistics, 33 (6), 574-616.


Derkarabetian S, M Burns, J Starrett & M Hedin.. 2016. Population genomic evidence for multiple Pliocene refugia in a montane-restricted harvestman (Arachnida, Opiliones, Sclerobunus robustus) from the southwestern United States. Molecular Ecology 25, 4611-4631.

DiDomenico A & M Hedin. 2016. New species in the Sitalcina sura species group (Opiliones, Laniatores, Phalangodidae), with evidence for a biogeographic link between California desert canyons and Arizona sky islands. ZooKeys 586: 1-36. doi: 10.3897/zookeys.586.7832

Emata KN & M Hedin. 2016. From the mountains to the coast and back again: Ancient biogeography in a radiation of short-range endemic harvestmen from California. Molecular Phylogenetics & Evolution 98:233-243.

Garrison NL, J Rodriguez, I Agnarsson, JA Coddington, CE Griswold, CA Hamilton, M Hedin, KM Kocot, JM Ledford & JE Bond. 2016. Spider Phylogenomics: Untangling the Spider Tree of Life. PeerJ 4:e1719; DOI 10.7717/peerj.1719

Richart CH, CY Hayashi & M Hedin. 2016. Phylogenomic analyses resolve an ancient trichotomy at the base of Ischyropsalidoidea (Arachnida, Opiliones) despite high levels of gene tree conflict and unequal minority resolution frequencies. Molecular Phylogenetics & Evolution 95:171-182.

Starrett J, S Derkarabetian, CH Richart, A Carbero & M Hedin.. 2016. A new monster from southwest Oregon forests: Cryptomaster behemoth sp. nov. (Opiliones, Laniatores, Travunioidea). ZooKeys 555: 11-35. doi: 10.3897/zookeys.555.6274

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